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Like Blaylock said it looks like you have a blockage in your block or rad possibly that is causing the restriction in the system which is creating extra head pressure against the pump. By adding more fluid to the reservoir it is allowing the extra pressure from the fluid in the reservoir to help overcome the head pressure from the restriction. I guess there is the possibility that the system is airlocked but I doubt this is the cause. You could try tipping the system on its side while filling/priming to see if you can get the air to move if it is airlocked. MY money is on a restriction somewhere thoguh especially if the system was not flushed prior to assembaly.
what kind of fittings are you using on your cpu block also what kind of cpu block is it? make sure the fittings attached to your cpu block arent bottomed out. some fittings have longer threaded parts. or if you use adapters etc they may also have threaded parts that bottom out.
One thing you could test is blowing through the block and radiator to see if it is hard to blow through one or the other. Did you check the in/out ports on the cpu block to make sure coolant flow was in the correct direction?
what kind of fittings are you using on your cpu block also what kind of cpu block is it? make sure the fittings attached to your cpu block arent bottomed out. some fittings have longer threaded parts. or if you use adapters etc they may also have threaded parts that bottom out.
Very glad you got your system up and running it looks nice !
What kind of power connector does your Pump have that you have yo make one ? isnt it a PWM 4 pin or a reg 3 pin fan connector ? If it is one of these you can probably plug it into your motherboard and make use of temp vs speed control for your pump .